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His main interest was demography , which possibly influenced his professional choice as a statistician at Metropolitan Life Insurance. One of Lotka's earliest publications, in , proposed a solution to Ronald Ross 's second malaria model.

In , he published a thorough five-part analysis and extension of both Ross's malaria models. The fourth part in the series, co-authored by F.

Sharpe, modeled the time lag for pathogen incubation. He is also known for his energetics perspective on evolution. Lotka proposed that natural selection was, at its root, a struggle among organisms for available energy; Lotka's principle states that organisms that survive and prosper are those that capture and use energy more efficiently than its competitors.

Lotka extended his energetics framework to human society. In particular, he suggested that the shift in reliance from solar energy to nonrenewable energy would pose unique and fundamental challenges to society.

These theories made Lotka an important forerunner to the development of biophysical economics and ecological economics , advanced by Frederick Soddy , Howard Odum and others.

His first book summarizes his previous work and organizes his ideas of unity and universality of physical laws, making his works accessible to other scientists.

Although the book covered a large amount of topics, from energetics of evolution see below to the physical nature of consciousness, the author is primarily known today for the Lotka—Volterra equation of population dynamics.

His earlier work was centered on energetics and applications of thermodynamics in life sciences. Lotka proposed the theory that the Darwinian concept of natural selection could be quantified as a physical law.

The law that he proposed was that the selective principle of evolution was one which favoured the maximum useful energy flow transformation. The general systems ecologist Howard T.

Odum later applied Lotka's proposal as a central guiding feature of his work in ecosystem ecology. Odum called Lotka's law the maximum power principle.

Lotka's work in mathematical demography began in with the publication of articles in the journal Science and American Journal of Science.

Within the field of bibliometrics, particularly that part devoted to studying scientific publications, Lotka is noted for contributing " Lotka's law ".

The law, which Lotka discovered, relates to the productivity of scientists. As noted by W.

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